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581.526.427 (HEDYSARUM GRANDIFLORUM PALL.) .. E.A. Averinova The ...

, 2014. Bulletin of Bryansk dpt. of RBS, 2014.

1(3). . 3747. N 1(3). P. 3747.

581.526.427

(HEDYSARUM GRANDIFLORUM PALL.)

.. E.A. Averinova The communities with Hedysarum grandiflorum Pall. within the territory of the Middle Russian upland . . .. , 241036, , . , . , 14. .: +7 (4832) 66-68-34, e-mail: elena_averi@mail.ru . (Hedysarum grandiflorum Pall.) . . Cephalario uralensisHedysaretum grandiflori ass. nov. Centaureo carbonataeKoelerion talievii Romaschenko, Didukh et V. Sl. 1996 ( HelianthemoThymetea). Hedysaro grandiflori Centauretum sumensis ass. nov. Carici humilisThymion calcarei all. nov. ( Festuco Brometea), .

: , Carici humilisThymion calcarei all. nov., HelianthemoThymetea, Festuco Brometea, , , , , .



Abstract. In the article the syntaxonomy of the communities with Hedysarum grandiflorum Pall. within the territory of the Middle Russian upland is worked out. Two new associations and one community without syntaxonomical status are described. The association Cephalario uralensisHedysaretum grandiflori ass. nov. is referred to the alliance Centaureo carbonataeKoelerion talievii Romaschenko, Didukh et V. Sl. 1996 (class HelianthemoThymetea). The association Hedysaro grandifloriCentauretum sumensis ass. nov. is referred to the new alliance Carici humilisThymion calcarei all. nov. (class FestucoBrometea) which includes the calciphytic thyme steppes of the Middle Russian upland.

Keywords: syntaxonomy, Carici humilisThymion calcarei all. nov., HelianthemoThymetea, FestucoBrometea, calciphilous species, calciphytic communities, thyme steppes, cretaceous outcrops, Middle Russian upland.

Hedysarum grandiflorum Pall. , , , - , , , .

(, ) ( ..., 1987). , , , , . : , , , , , , , , , , ( , 2008). (, 1917), ( , 2002; , 1998). , (, 2005).





(2008) 3 . , . , , , , , .

(2004) VI - . 9 . (2011) II . 10 .

20112012 . 11 100 2. , .. ( ., 1996).

- (Westhoff, Maarel, 1978) TURBOVEG (Hennekens, 1995) JUICE (Tich et al., 2011), , Statistica 6.0.

, (Kopek, Hejn, 1974). (Weber et al., 2000). .. (1995).

FestucoBrometea HelianthemoThymetea, . , . , 3 .. .

, , . .

FestucoBrometea Br.-Bl. et R. Tx. in Br.-Bl. 1949 Festucetalia valesiacae Br.-Bl. et R. Tx. ex Br.-Bl. 1949 Carici humilisThymion calcarei all. nov. hoc loco . Hedysaro grandifloriCentauretum sumensis ass. nov. hoc loco C. H. g.C. s. adonidetosum vernalis subass. nov. hoc loco C. H. g.C. s. typicum subass. nov. hoc loco HelianthemoThymetea Romaschenko, Didukh et V. Sl. 1996 Thymo cretaceiHyssopetalia cretacei Didukh 1989 Centaureo carbonataeKoelerion talievii Romaschenko, Didukh et V. Sl. 1996 . Cephalario uralensisHedysaretum grandiflori ass. nov. hoc loco Hedysarum grandiflorum [FestucoBrometea/HelianthemoThymetea] . Hedysaro grandifloriCentauretum sumensis ass. nov. hoc loco (. 1, . 17; (holotypus) . 6).

. : Allium flavescens, A. inaequale, Centaurea orientalis, C. sumensis, Hedysarum grandiflorum, Helianthemum canum, Melampyrum argyrocomum. - Carex humilis. Thymus calcareus, Onosma simplicissima, Stipa pennata, S. capillata, Gypsophila altissima. - 45 80 % ( 55 %). ( ) 5 25 . , Cephalaria uralensis Gypsophila altissima 50 . Abietinella abietina 25 %. 31 52 100 2 ( 41).

. ( 2 7, 10) (-, ) . , .

.

. H. g.C. s. adonidetosum vernalis subass. nov. hoc loco (. 1, . 14; (holotypus) . 2).

: Adonis vernalis, Anthericum ramosum, Astragalus onobrychis, Helianthemum nummularium, Hypericum elegans, Nonea pulla, Onobrychis arenaria, Seseli annuum. . . - Adonis vernalis .

. H. g.C. s. typicum subass. nov. hoc loco (. 1, . 57; (holotypus) . 6) . ( 5) Hedysaro ucrainiciArtemisietum hololeucae Averinova 2011 violetosum ambiguae subass. prov. (, 2011).

   

. * .

: Adonis wolgensis 13 (+), Agrimonia eupatoria 3 (r), 14 (+), Agropyron cristatum 7 (+), Ajuga chia 1 (r), A. glabra 13 (+), A. laxmannii 1 (r), Allium rotundum 13 (+), A. sphaerocephalum 1 (r), 10 (+), Androsace elongata, A. septentrionalis 14 (+), Anemone sylvestris 5 (r), 14 (+), Artemisia austriaca 7 (r), Astragalus danicus 14 (+), A. ucrainicus 12 (+), A. varius 5 (r), Bromopsis inermis 14 (+), Calamagrostis epigeios 1, 14 (+), Chaenorhinum minus 10 (r), Chamaecytisus ruthenicus 3 (r), 5 (+), Cichorium intybus 6 (r), Clausia aprica 2 (r), Cleistogenes squarrosa 13 (+), Collema sp. 6 (+), Crambe tataria 12 (+), Cuscuta europaea 14 (1), Elytrigia intermedia 1 (+), Ephedra distachya 7 (r), Eremogone micradenia 7 (+), Erigeron acris 3 (r), Eryngium campestre

14 (+), Euphorbia stepposa 5, 14 (+), E. uralensis 13 (+), E. virgata 14 (+), Euphrasia sp. 6, 9 (+), E. tatrae 14 (+), Filipendula vulgaris 3 (r), Fragaria viridis 14 (+), Galatella linosyris 12 (+), G. villosa, Galium humifusum 13 (+), G. tinctorium 2 (r), 12 (+), G. verum 3 (+), Gentiana cruciata 14 (+), Helictotrichon schellianum 2 (+), Hieracium sp. 2, 5 (+), Iris sp. 7 (r), Lactuca tatarica 13 (+), Limonium tomentellum 12 (+), Linaria cretacea 14 (+), Lithospermum officinale 7 (r), Marrubium praecox 1 (r), 14 (+), Melilotus albus 7 (r), Odontites vulgaris 9, 14 (+), Oxytropis pilosa 3 (r), Paeonia tenuifolia 7 (r), 13 (2), Pinus sylvestris 6 (+), 10 (r), Plantago lanceolata 14 (+), P. media 5 (r), 14 (+), Poa angustifolia 14 (+), P. compressa 5 (r), 6 (+), P. pratensis 13 (+), Potentilla recta, Prunella vulgaris, Ranunculus polyanthemos 14 (+), Rosa sp. 10 (r), Salvia aethiopis, S. tesquicola 14 (+), Scutellaria supina 7 (r), Seseli libanotis 3, 5 (+), Silene artemisetorum 13 (+), Verbascum lychnitis 1, 7 (r), Veronica incana 14 (+), V. spicata 5 (+), Vinca herbacea 1, 13 (+), V. minor, Vincetoxicum rossicum 14 (+).

. . -, . : . 1, 3, 8-11 , -, , 4.08.2012, .. ; . 2 , 3.08.2012, .. ; . 14 , 25.08.1983, .. ; . 4, 6 .

, 1.08.2012, .. . -, , . : . 5 1 . , , 6.08.2011, . 7 1 . , , 9.08.2011, .. . -: . 12 0,5 . , . , 24.08.1983, .. . . . 13 -, , 1.08.1983, .. .

. Hedysaro grandifloriCentauretum sumensis ass. nov. Carici humilisThymion calcarei all. nov. hoc loco, Festucetalia valesiacae. (holotypus) Carici humilisThymetum calcarei Poluyanov 2009 : 53, 5660.

: Adonis vernalis, Androsace koso-poljanskii (dom.), Asperula cynanchica, Astragalus albicaulis, A. austriacus, Carex humilis (dom.), Centaurea sumensis, Festuca valesiaca, Helianthemum nummularium, Linum ucrainicum, Medicago falcata, Onosma simplicissima (dom.), Pimpinella tragium, Polygala cretacea, P. sibirica, Salvia verticillata, Stachys recta, Stipa pennata, Thymus calcareus (dom.), Viola ambigua. . 2. , Carici humilisThymion calcarei ( HelianthemoThymetea FestucoBrometea). , . .. ( ., 2009), . , HelianthemoThymetea , FestucoBrometea, .

   

Centaureo carbonataeKoelerion talievii .

Sileno supinaeArtemision hololeucae Didukh 1989. Artemisio hololeucaeHyssopion cretacei Romaschenko, Didukh et V. Sl. 1996 nomen superfluum.

: 1 Thymo cretaceiHyssopetum cretacei Didukh 1989; 2 Pimpinello titanophyllaeArtemisietum salsoloidis Didukh 1989; 3 Euphorbio cretophilaeJurinetum brachycephalae Didukh 1989; 4 Jurineo brachycephalaeHelianthemetum cretophilae Romaschenko, Didukh et V. Sl. 1996; 5 Onosmo tanaiticae Androsacietum kozo-poljanskii Romaschenko, Didukh et V. Sl. 1996; 6 Scrophulario cretaceiHelianthemetum cretacei Romaschenko, Didukh et V. Sl. 1996; 7 Sileno supinaeMatthioletum fragrantis Didukh 1989; 8 Artemisio hololeucaePolygaletum cretaceae Didukh 1989; 9 Artemisio nutantisPlantaginetum salsae Didukh 1989; 10 Hedysaro ucrainiciArtemisietum hololeucae Averinova 2011; 11 Jurineo brachycephalaeKoelerietum talievii Romaschenko, Didukh et V. Sl. 1996; 12 Gypsophilo oligospermaeCampanuletum sibiricae Romaschenko, Didukh et V.

Sl. 1996; 13 Cephalario uralensisHedysaretum grandiflori ass. nov.; 14 Astragalo albicaulisCephalarietum uralensis Averinova 2012; 15 Hedysaro grandifloriCentauretum sumensis adonidetosum vernalis subass. nov.; 16 H. g.C. s. typicum subass. nov.; 17 Androsacio kozo-poljanskiiCaricetum humilis Korotchenko et Didukh 1997; 18 Carici humilisThymetum calcarei typicum Poluyanov 2009; 19 C. h.T. c. androsacietosum koso-poljanskiae Poluyanov 2009; 20 Astero amelliPotentilletum humifusae Poluyanov in Poluyanov et Averinova 2012; 21 Asperulo cynanchicaeOnobrychidetum arenariae Averinova 2005; 22 Allio rotundiAstragaletum onobrychis (Averinova 2005) Poluyanov et Averinova 2012; 23 Gypsophilo altissimaeStipetum capillatae Poluyanov 2009; 24 Inulo ensifoliae Stipetum pennatae Poluyanov in Poluyanov et Averinova 2012; 25 Stachyo rectaeEchinopetum ruthenici Averinova 2010 ass. prov.; 26 Allio paniculatiGypsophiletum altissimae Averinova 2012 ass. prov.

( ): 13, 79 (ij, 1989); 46, 11, 12 ( ., 1996); 14 (, 2012); 17 (, i, 1997); 18, 19 (, 2009); 2024 (, 2010 ;

, , 2012); 25 (, 2010); 26 (, 2012 ).

CariciThymion - , HelianthemoThymetea ( ) FestucoBrometea (). . Thymus calcareus, Onosma simplicissima, Androsace koso-poljanskii, Astragalus albicaulis, Carex humilis. Stipa pennata, S. capillata. , . 10 80 %. Abietinella abietina.

4 : Hedysaro grandifloriCentauretum sumensis ass. nov., Astragalo albicaulisCephalarietum uralensis Averinova 2012, Centaureo carbonataeKoelerion talievii (, 2012), Carici humilisThymetum calcarei Poluyanov 2009 Androsacio kozo-poljanskiiCaricetum humilis Korotchenko et Didukh

1997. Centaureo carbonataeKoelerion talievii.

, CariciThymion Centaureo carbonataeKoelerion talievii, (. 2). HelianthemoThymetea ( ., 1996), Festuco Brometea (, i, 1997). (, , 2012) CentaureoKoelerion (. Carici humilisThymetum calcarei). CentaureoKoelerion FestucoBrometea .

, CariciThymion Bupleuro falcatiGypsophilenion altissimae Averinova 2005 (Festucion valesiacae), (, 2005, 2010, 2010 , 2012 ). CariciThymion HelianthemoThymetea, , .

CariciThymion (.). HelianthemoThymetea FestucoBrometea. HelianthemoThymetea ( III) , . FestucoBrometea 2 : I BupleuroGypsophilenion, II CariciThymion.

Ward`s

   

. .

: I BupleuroGypsophilenion (Festucion valesiacae); II Carici humilisThymion calcarei; III HelianthemoThymetea.

: 1 Cephalario uralensisHedysaretum grandiflori; 2 Hedysaro grandifloriCentauretum sumensis adonidetosum vernalis; 3 H. g.C. s. typicum; 4 Astragalo albicaulisCephalarietum uralensis; 5 Jurineo brachycephalaeKoelerietum talievii; 6 Gypsophilo oligospermaeCampanuletum sibiricae; 7 Onosmo tanaiticae Androsacietum kozo-poljanskii; 8 Scrophulario cretaceiHelianthemetum cretacei; 9 Sileno supinaeMatthioletum fragrantis; 10 Artemisio hololeucaePolygaletum cretaceae; 11 Artemisio nutantisPlantaginetum salsae; 12 Thymo cretaceiHyssopetum cretacei; 13 Pimpinello titanophyllaeArtemisietum salsoloidis; 14 Euphorbio cretophilaeJurinetum brachycephalae; 15 Jurineo brachycephalaeHelianthemetum cretophilae; 16 Hedysaro ucrainiciArtemisietum hololeucae Averinova 2011 violetosum ambiguae subass. prov.; 17 H. u.A. h. typicum; 18 Polygalo sibiricaeHyssopetum cretacei Poluyanov in Poluyanov et Averinova 2012; 19 Androsacio kozo-poljanskii Caricetum humilis; 20 Bupleuro falcataeStipetum capillatae Romaschenko, Didukh et V. Sl. 1996; 21 Carici humilisThymetum calcarei typicum; 22 C. h.T. c. androsacietosum koso-poljanskiae; 23 Teucrio polii Euphorbietum stepposae Averinova 2011; 24 Astragalo dasyanthiChamaecytisetum austriaci Averinova nom. ined.;

25 Astero amelliPotentilletum humifusae; 26 Asperulo cynanchicaeOnobrychidetum arenariae; 27 Allio rotundiAstragaletum onobrychis; 28 Gypsophilo altissimaeStipetum capillatae; 29 Inulo ensifoliaeStipetum pennatae; 30 Stachyo rectaeEchinopetum ruthenici; 31 Allio paniculatiGypsophiletum altissimae.

. Cephalario uralensisHedysaretum grandiflori ass. nov. hoc loco (. 1, . 811;

(holotypus) . 8).

. : Cephalaria uralensis, Diplotaxis cretacea, Hedysarum grandiflorum (dom.), Jurinea arachnoidea, Thesium arvense, Thymelaea passerina. . arex humilis , Thymus calcareus. - 3 . 3 Thymus calcareus. 1015 Hedysarum grandiflorum, Cephalaria uralensis, Onosma simplicissima. 4070 Hedysarum grandiflorum, Cephalaria uralensis, Echinops ruthenicus. -- . - . , . 4065 % ( 50 %). .

23 33 100 2 ( 27).

. (520) , , - . .

. Centaureo carbonataeKoelerion talievii ( HelianthemoThymetea). .

Hedysarum grandiflorum [FestucoBrometea/Helianthemo Thymetea] (. 1, . 1214).

: Adonis vernalis, Allium sphaeropodum, Centaurea ruthenica, Hedysarum grandiflorum, Linum flavum, L. perenne, Pimpinella saxifraga, Phlomis pungens, Silene cretacea, Stipa lessingiana. . (Carex humilis, Stipa capillata), HelianthemoThymetea (Artemisia hololeuca, A. nutans, Astragalus albicaulis, Hyssopus cretaceus, Onosma simplicissima, Thymus calcareus). 60 65 %, . 37 74 100 2. .. ( ., 1996) .

.. .. . .. .

.. ( ) // . 2005. 7. . 3949.

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